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Hall, H. G. 1992a. DNA studies reveal processes involved in the spread of New World African honeybees. Insect Behavioral Ecology 75(1): 51-59.

Hall, H. G. 1992b. Suspected African honeybee colonies in Florida tested for identifying DNA makers. Fla. Entomol. 75(2): 257-266.

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Hammer, M. & R. Menzel, 1998. Multiple sites of associative odor learning as revealed by local brain microinjections of octopamine in honeybees. Learning & Memory 5: 146-156.

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Harbo, J. R. 1979a. Egg hatch of honey bees fertilized with frozen spermatozoa. Annals of the Entomological Society of America 72(4): 516-518.

Harbo, J. R. 1979b. Storage of honeybee spermatozoa at -196 degrees Celsius. Journal of Apicultural Research 18(1): 57-63.

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Harbo, J. R. 1980. Mosaic male honeybees produced by queens inseminated with frozen spermatozoa. Journal of Heredity 71(6): 435-436.

Harbo, J. R. 1981. Viability of honeybee eggs from progeny of frozen spermatozoa. Annals of the Entomological Society of America 74(5): 482-486.

Harbo, J. R. 1983. Effect of population size on worker survival and honey loss in broodless colonies of honey bees, Apis mellifera L. (Hymenoptera: Apidae). Environmental Entomology 12(5): [page no.?].

Harbo, J. R. 1983. Survival of honey bee (Hymenoptera:Apidae) spermatozoa after two years in liquid nitrogen (-196 degrees C). Annals of the Entomological Society of America 76(5): 890-891.

Harbo, J. R. 1985a. Instrumental insemination of queen bees, 1985. 1. American Bee Journal 125(3): 197-202.

Harbo, J. R. 1985b. Instrumental insemination of queen bees, 1985: 2. American Bee Journal 125(4): 282-283, 285-287 (Concluded).

Harbo, J. R. 1986a. Effect of population size on brood production, worker survival and honey gain in colonies of honeybees. Journal of Apicultural Research 25(1): 22-29.

Harbo, J. R. 1986b. Oviposition rates of instrumentally inseminated and naturally mated queen honey bees (Hymenoptera: Apidae). Annals of the Entomological Society of America 79(1): 112-115.

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Harbo, J. R. 1988b. Sperm competition. American Bee Journal 128(12): 803-804.

Harbo, J. R. 1990. Artificial mixing of spermatozoa from honeybees and evidence for sperm competition. Journal of Apicultural Research 29(3): 151-158.

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Harbo, J. R. 1992. Breeding honey bees (Hymenoptera: Apidae) for more rapid development of larvae and pupae. Journal of Economic Entomology 85(6): 2125-2130.

Harbo, J. R. 1993a. Field and laboratory tests that associate heat with mortality of tracheal mites. Journal of Apicultural Research 32(3/4): 159-165.

Harbo, J. R. 1993b. Worker-bee crowding affects brood production, honey production, and longevity of honey bees (Hymenoptera: Apidae). Journal of Economic Entomology 86(6): 1672-1678.

Harbo, J. R. 1993c. Effect of brood rearing on honey consumption and the survival of worker honey bees. Journal of Apicultural Research 32(1): 11-17.

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Harbo, J. R. & J. W. Harris, 1999a. Heritability in honey bees (Hymenoptera: Apidae) of characteristics associated with resistance to Varroa jacobsoni (Mesostigmata: Varroidae). Journal of Economic Entomology 92(2): 261-265.

Harbo, J. & J. Harris, 1999b. Selecting honey bees for resistance to Varroa jacobsoni. Apidologie 30: 183-196.

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Harbo J. R. & J. W. Harris, 2000. Using free-mated queens to introduce genes for varroa resistance into a population of honey bees. American Bee Journal 140: 904-905.

Harbo, J. R. & J. W. Harris, 2001a. Resistance to Varroa destructor (Mesostigmata: Varroidae) when mite-resistant queen honey bees (Hymenoptera: Apidae) were free-mated with unselected drones. Journal of Economic Entomology 94(6): 1319-1323.

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Harbo, J. R. & J. L. Williams, 1987. Effect of above-freezing temperatures on temporary storage of honeybee spermatozoa. Journal of Apicultural Research 26(1): 53-55.

Harbo, J. R. & J. L. Zuhlke, 1988. Populations of Varroa jacobsoni in a Florida apiary. American Bee Journal 128(11): 737-739.

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Harris, J. W., & J. R. Harbo, 2000. Changes in reproduction of Varroa destructor after honey bee queens were exchanged between resistant and susceptible colonies. Apidologie 31(6): 689-699.

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Harris, J. W. 2001. A technique for marking individual varroa mites. Journal of Apicultural Research 40(1): 35-37.

Harris, J. W. & J. R. Harbo, 1988. Breeding from worker honey bees. American Bee Journal 128(12): 804.

Harris J. W. & J. R. Harbo, 1989. Ovary development of worker bees when caged with workers from different stocks. American Bee Journal 128: 816.

Harris, J. W. & J. R. Harbo, 1990. Suppression of ovary development of worker honeybees by association with workers treated with carbon dioxide. Journal of Apicultural Research 29(4): 187-193.

Harris, J. W. & J. R. Harbo, 1991. Producing eggs from a single worker honey bee (Hymenoptera: Apidae). Journal of Economic Entomology 84(3): 818-824.

Harris J. W. & J. R. Harbo, 1998. Low sperm counts and delayed oviposition of mites in colonies of honey bees that are resistant to Varroa jacobsoni. American Bee Journal 138: 296.

Harris, J. W. & J. R. Harbo, 1999. Low sperm counts and reduced fecundity of mites in colonies of honey bees (Hymenoptera: Apidae) resistant to Varroa jacobsoni (Mesostigmata: Varroidae). Journal of Economic Entomology 92(1): 83-90.

Harris J. W. & J. R. Harbo, 2000. Changes in reproduction of Varroa destructor after honey bee queens were exchanged between resistant and susceptible colonies. Apidologie 31: 689-699.

Harris J. W. & J. R. Harbo, 2001. Natural and suppressed reproduction of varroa mites. Bee Culture 129: 34-38.

Harris J. W., J. R. Harbo & J. P. Woodring, 1999. A comparison of hemolymph proteins in honey bee larvae that are resistant and susceptible to Varroa jacobsoni. American Bee Journal 139: 312.

Harris J. [W.], T. Rinderer, V. Kuznetsov, R. Danka, G. Delatte, L. de Guzman and J. Villa, 2002. Imported Russian Honey Bees: Quarantine and Initial Selection for Varroa Resistance. American Bee Journal 142: 591-596.

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Hennig, W. 1969. Die Stammesgeschichte der Insekten. 436 pages. W. Kramer, Frankfurt am Main.

Hennig, W. 1974. Kritische Bemerkungen zur Frage “Cladistic analysis or cladistic classification ?” Z. Zool. Syst. Evolutionsforsch., Hamburg 12: 279-294.

Henschel, G. 1988. Megachile villosa. Ein biologischer Beitrag. Entomologische Nachrichten (Karsch: Berlin) 14: 321-323.

Hensel, A. 1870. Prosopis gerstaeckeri, eine muthmasslich neue Bienen Art. Beitr. Entomol. Zeit. xiv: 185-190.

Hensen, R. V. 1987. Hylaeus (Metylaeus) mahafaly sp.n., a new Malagasy bee. Entomologische Berichten 47: 152-154.

Henslow, G. 1867. Note on the structure of Medicago sativa, as apparently affording facilities for the intercrossing of distinct flowers. J. Linnean Soc. 9: 327-329.

Henslow, G. 1888. The origin of floral structure through insect and other agencies. International Scientific Series, 63 London: 1888.

Heon, S. 2000. The mite Varroa, a menace to the bees in our hives. Nouv'Ailes 10(2): 1-4.

Hepburn, H.R. 1978a. The bee louse. South African Bee Journal 50(6): 11-12.

Hepburn, H. R. 1978b. Some honeybee research activities at the University of the Witwatersrand. South African Bee Journal 50(5): 11-14.

Hepurn, H. R. 1979a. Braula coeca. South African Bee Journal 50(6): 11-12.

Hepurn, H. R. 1979b. Pollination versus honey: the beekeeper’s seesaw. South African Bee Journal 51(2): 3-7.

Hepburn, H. R. 1980. How honeybees measure their pollen loads. South African Bee Journal 52(3): 11-13.

Hepurn, H. R. 1981a. Efficacy of pollen traps. South African Bee Journal 53(5): 8-9.

Hepurn, H. R. 1981b. Fruit tree pollination. South African Bee Journal 53(5): 14-15.

Hepurn, H. R. 1981c. Is the Langstroth brood chamber deep enough for African bees? South African Bee Journal 53(4): 9-11.

Hepurn, H. R. 1982a. Comb production and the new foundation. South African Bee Journal 54: 56-62.

Hepurn, H. R. 1982b. The bee scorpion. South African Bee Journal 54(2): 39-41.

Hepburn, H. R. 1983a. Comb construction by the African honey bee, Apis mellifera adansonii. Journal of the Entomological Society of South Africa 46(1): 87-101.

Hepurn, H. R. 1983b. Single versus double brood chambers. South African Bee Journal 55: 11-13.

Hepurn, H. R. 1986. Honeybees and Wax: an Experimental Natural History. Springer-Verlag, Berlin 205pp.

Hepburn, H. R. 1988. Absconding in the African honey bee: The queen, engorgement and wax secretion. Journal of Apicultural Research 27(2): 95-102.

Hepurn, H. R. 1989. Capensis in the Eastern Cape. South African Bee Journal 61: 87-89.

Hepurn, H. R. 1990a. The first report on the biology of the Cape honeybee. South African Bee Journal 62: 128-132.

Hepurn, H. R. 1990b. What and where is the Cape bee: an interim report. Pp. 74-83. In: Anderson, R. H. & B. Buys (Eds.), Bees and Beekeeping in Southern Africa. Apimondia, Stellenbosch.

Hepburn, H. R. 1991a. Incidence of the tachinid bee fly, Rondaniooestrus apivorus, in southern Africa. Journal of the Entomological Society of Southern Africa 54(1): 84-85.

Hepurn, H. R. 1991b. Some apicultural puzzles. South African Bee Journal 63: 13-15.

Hepurn, H. R. 1991c. The unique Cape bee and some apicultural puzzles. Beeline 28: 12-15.

Hepurn, H. R. 1992a. “Follow-flow” brood rearing in capensis and scutellata. South African Bee Journal 64: 104-108.

Hepurn, H. R. 1992b. Drongos: honey guides of the eastern Cape. South African Bee Journal 64: 40-41.

Hepurn, H. R. 1992c. Pheromonal and ovarial development covary in Cape worker honeybees, Apis mellifera capensis. Naturwissenschaften 79: 523-524.

Hepurn, H. R. 1992d. The biology of queen production in the Cape honeybee. South African Bee Journal 64: 132-138.

Hepurn, H. R. 1993a. Heuning tydens die Tweede Vryheidsoorlog. South African Bee Journal 65: 10-12.

Hepurn, H. R. 1993b. Swarming, absconding and migration in southern African bees. South African Bee Journal 65: 61-66.

Hepburn, H. R. 1994. Reproductive cycling and hierarchical competition in Cape honeybees, Apis mellifera capensis Esch. Apidologie 25: 38-48.

Hepurn, H. R. 1995. Flight and foraging in capensis and scutellata. South African Bee Journal 67: 38-42.

Hepurn, H. R. 1998. Reciprocal interactions between honeybees and combs in the integration of some colony functions in Apis mellifera. Apidologie 29: 47-66.

Hepurn, H. R. 2000. Honey bee (Apis mellifera) classification and the confounding effects of trinomial nomenclature. Anais do Encontro sobre Abelhas 4: 188-196.

Hepburn, H. R. 2001. The enigmatic Cape honey bee, Apis mellifera capensis. Bee World 82(4): 181-191. also in Honeybee Science 22: 49-56. [In Japanese].

Hepurn, H. R. 2002. Apis mellifera capensis: concept and reality. South African Bee Journal 74(2): 37-39, 53-62.

Hepurn, H. R. & M. H. Allsopp, 1994. Reproductive conflict between honeybees: usurpation of Apis mellifera scutellata colonies by Apis mellifera capensis. South African Journal of Science 90: 247-249.

Hepurn, H. R., E. Armstrong & S. P. Kurstjens, 1983. The ductility of native beeswax is optimally related to honeybee colony temperature. South African Journal of Science 79: 416-417.

Hepurn, H. R., R. T. F. Bernard, B. C. Davidson, W. J. Muller, P. Lloyd, S. P. Kurstjens & S. L. Vincent, 1991. Synthesis and secretion of beeswax in honeybees. Apidologie 22: 21-36.

Hepburn, H. R., R. C. Cantrill, P. R. Thompson & E. Kennedi, 1979. Metabolism of carbohydrate, lipid and protein during development of sealed worker brood of the African honeybee Apis mellifera adansonii. Journal of Apicultural Research 18(1): 30-35.

Hepburn, H. R. & H. D. Chandler, 1975. A new classification for arthropod cuticles. South African Journal of Science 71(10): 311-312.

Hepburn, H. R. & H. D. Chandler, 1976a. Material properties of arthropod cuticles: the arthrodial membranes. Journal of Comparative Physiology 109(2): 177-198.

Hepburn, H. R. & H. D. Chandler, 1976b. Mechanical hysteresis of insect cuticles. Journal of Insect Physiology 22(2): 221-226.

Hepburn, H. R., H. D. Chandler & M. R. Davidoff, 1979. Extensometric properties of insect fibroins: the green lacewing cross-beta, honeybee alpha-helical and greater waxmoth parallel-beta conformations [Chrysopa carnea, Apis mellifera adansonii, Galleria mellonella]. Insect Biochemistry 9(1): 69-77.

Hepburn, H. R. & R. M. Crewe, 1990. Defining the Cape South Africa honey bee reproductive traits of queenless workers. South African Journal of Science 86: 524-527.

Hepburn, H. R. & R. M. Crewe, 1991a. Portrait of the Cape honeybee, Apis mellifera capensis. Apidologie 22(6): 567-580.

Hepurn, H. R. & R. M. Crewe, 1991b. Geography of the Cape honeybee based on laying worker performance. South African Bee Journal 63: 51-59.

Hepburn, H. R. & S. G. Guye, 1993. An annotated bibliography of the Cape honeybee, Apis mellifera capensis Eschscholtz (Hymenoptera: Apidae). African Entomology 1(2): 235-252.

Hepburn, H. R. & C. Hepburn, 2005. Bibliography of Apis florae. Apidologie 36(3): 377-378.

Hepburn, H. R., J. J. Hugo, D. Mitchell, M. J. M. Nijland & A. G. Scrimgeour, 1984. On the energetic costs of wax production by the African honeybee, Apis mellifera adansonii. South African Journal of Science 80(8): 363-368.

Hepurn, H. R. & Guillarmod A. Jacot, 1991. The Cape honeybee and the fynbos biome. South African Journal of Science 87: 70-73.

Hepurn, H. R., G. E. Jones & R. Kirby, 1994. Introgression between Apis mellifera capensis Escholtz and Apis mellifera scutellata Lepeletier: the sting pheromones. Apidologie 25: 557-565.

Hepburn, H. R. & S. P. Kurstjens, 1984. On the strength of propolis (bee glue). Naturwissenschaften 71(11): 591-592.

Hepburn, H. R. & S. P. Kurstjens, 1988. The combs of the honeybee as composite materials. Apidologie 19: 25-36.

Hepburn, H. R. & P. C. Magnuson, 1988. Nectar storage in relation to wax secretion by honeybees. Journal of Apicultural Research 27(2): 90-92.

Hepburn, H. R., P. Magnuson, L. Herbert & L. A. Whiffler, 1991. The development of laying workers in field collected colonies of the Cape honeybee. J. Apic. Res. 30: 107-112.

Hepurn, H. R. & G. Mitchell, 1981. Milk and Honey, Essays in Agriculture. Elsevier/North Holland, Amsterdam 108 pp.

Hepurn, H. R. & W. J. Muller, 1988. Wax secretion in honeybees. Thermal integrity of the festoon. Naturwissenschaften 75: 628-629.

Hepurn, H. R., R. J. C. Nefdt & L. A. Whiffler, 1988. Queen loss in the Cape honeybee: the interactions of brood, laying workers (false queens) and queen cells. South African Journal of Science 84: 778-780.

Hepburn, H.R. & S. E. Radloff, 1995. First approximation to a phenology of the honeybees (Apis mellifera) and flora of Africa. Oecologia 101(3): 265-273.

Hepurn, H. R. & S. E. Radloff, 1996a. Beeswax exports and rainfall in the savanna woodlands of east central Africa. Apidologie 27: 473-478.

Hepurn, H. R. & S. E. Radloff, 1996b. Population structure of the honeybees along a transect from the Sahara to the Pyrenees. Apidologie 27: 35-45.

Hepurn, H. R. & S. E. Radloff, 1997. Biogeographical correlates of population variance in the honeybees (Apis mellifera) of Africa. Apidologie 28: 243-258.

Hepurn, H. R. & S. E. Radloff, 1998. Honeybees of Africa. Springer-Verlag, Berlin 370 pp.

Hepburn, H. R. & S. E. Radloff, 2002. Apis mellifera capensis: an essay on the subspecific classification of honeybees. Apidologie 33(2): 105-128.

Hepburn, H. R., S. E. Radloff & S. Fuchs, 1998a. Population structure and the interface between Apis mellifera capensis and Apis mellifera scutellata. Apidologie 29: 333-346.

Hepurn, H. R., S. E. Radloff & S. Fuchs, 1998b. Ecological and morphological differentiation of the honeybees, Apis mellifera Linnaeus (Hymenoptera: Apidae) of west Africa. African Entomology 6: 17-23.

Hepburn, H. R., S. E. Radloff & S. Fuchs, 1999. Flight machinery dimensions of honeybees, Apis mellifera. Journal of comparative physiology. B. Biochemical, systemic, and environmental Physiology 169(2): 107-112.

Hepurn, H. R., S. E. Radloff, S. Fuchs, G. W. Otis, S. Hadisoesilo & A. C. Tilde, 2003. Mutlivariate morphometric analysis of the Apis cerana populations of oceanic southern Asia. Apidologie [In Press].

Hepburn, H. R., S. E. Radloff & S. Oghiakhe, 2000. Mountain honeybees of Africa. Apidologie 31(2): 205-222.

Hepburn, H. R., S. E. Radloff, G. W. Otis, S. Fuchs, L. R. Verma, T. Ken, T. Chaiyawong, G. Tahmasebi, R. Ebadi & S. Wongsiri, 2005. Apis florea: morphometrics, classification and biogeography. Apidologie 36(3): 359-376.

Hepburn, H. R., S. E. Radloff, G. R. Steele & R. E. Brown, 1998. Dimensional aspects of flight in the honey bees of Africa. Journal of Apicultural Research 37(3): 147-154.

Hepburn, H. R., S. E. Radloff, S. Verma & L. R. Verma, 2001. Morphometric analysis of Apis cerana populations in the southern Himalayan region. Apidologie 32: 1-13.

Hepurn, H. R., S. L. Reece, P. Neumann, R F. A. Moritz & S. E. Radloff, 1999. Absconding in honeybees (Apis mellifera) in relation to queen status and mode of worker reproduction. Insectes Sociaux 46: 323-326.

Hepburn, H. R. & J. R. Rigby, 1981a. Strength of beeswax foundation in relation to cell orientation. Journal of Apicultural Research 20(4): 234-238.

Hepurn, H. R. & J. R. Rigby, 1981b. Beeswax candles from a silicone rubber mould. South African Bee Journal 53(4): 11-13.

Hepburn, H. R. & D. C. Roberts, 1975. Stiffness and tanning of sclerites. J. Insect Physiol. 21(11): 1741-1746.

Hepburn, H. R. & G. N. Sinclair, 1977. On the occurrence of a fungus disease in the compound eyes of the African drone. South African Bee Journal 49(6): 2-4.

Hepburn, H. R., D. R. Smith, S. E. Radloff & G. W. Otis, 2001. Infraspecific categories of Apis cerana: morphometric, allozymal and mtDNA diversity. Apidologie 32(1): 3-23.

Hepburn, H. R., M. H. Villet, G. E. Jones, A. R. Carter, V. I. Simon & W. Coetzer, 1993. Winter absconding as a dispersal mechanism of the Cape honeybee. South African Journal of Science 89(6): 294-297.

Hepurn, H. R. & L. A. Whiffler, 1988. Dwarfs and queenballs in swarms of the Cape honeybee. South African Journal of Science 84: 851-852.

Hepurn, H. R. & L. A. Whiffler, 1991. Construction defects define pattern and method in comb building by honeybees. Apidologie 22: 381-388.

Hepburn, H. R., P. B. Wilson, H. D. Chandler & P. R. Thompson, 1978. Fractography of insect cuticle. Journal of Entomological Society of South Africa 41(1): 51-56.

Hepurn, H. R., C. Youthed, P. Illgner, S. E. Radloff & R. E. Brown, 1998. Production of aerodynamic power in mountain honeybees (Apis mellifera). Naturwissenschaften 85: 389-390.

Heppner, J. B. & D. H. Habeck, 1977. Insects associated with Polygonum (Polygonaceae) in north central Florida. II. Insects other than Lepidoptera. Fla. Entomol. 60: 167-170.

Heras, C. & S. F. Gayubo, 1989. Bee fauna from the province of Zamora [Spain]: II. Megachilidae (Hymenoptera, Apoidea). Bol. Asoc. Esp. Entomol. 13: 237-250.

Heraty, J. M., J. B. Woolley & D. C. Darling, 1994. Phylogenetic implications of the mesofurca and mesopostnotum in Hymenoptera. Journal of Hymenoptera Research 3: 241-277.

Herbers, J. M. 1979. The evolution of sex-ratio strategies in hymenopteran societies. American Naturalist 114: 818-834.

Herbert, E. W. Jr. 1979a. A new ash mixture for honeybees maintained on a synthetic diet. Journal of Apicultural Research 18(2): 144-147.

Herbert, E. W. Jr. 1979b. Brood rearing by small caged honeybee colonies fed whey-yeast pollen substitutes. Journal of Apicultural Research 18(1): 43-46.

Herbert, E. W., W. E. Bickley & H. Shimanuki, 1970. The brood-rearing capability of caged honey bees fed dandelion and mixed pollen diets. Journal of Economic Entomology 63(1): 215-218.

Herbert, E. W. Jr., W. A. Bruce & H. Shimanuki, 1988. Control of Varroa jacobsoni on honey bees in packages using Apistan. American Bee Journal 128(9): 615-616.

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