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El Pofesor Francis Ratnieks es el director del Apicultural and Social Insect Laboratory del Department of Animal and Plant Sciences, Sheffield University, Western Bank, Sheffield S10 2TN, Reino Unido. tel: 0114 2220070 (office), 01142 220147 (Lab. office), 01142220149 (Lab/postdocs/students), 0114 2220002 (fax). mailto:F.Ratnieks@Sheffield.ac.uk

publications in refereed journals and books (1987-2004) Editar

Submitted for publication Editar

Chaline, N., Sandoz, J. C., Martin, S. J., Ratnieks, F. L. W., Jones, G. R. Learning and discrimination of individual cuticular hydrocarbons by honey bees (Apis mellifera). (submitted)

Jensen, A. B., Palmer, K. A., Chaline, N., Raine. N. E., Tofilski, A., Pedersen, B. V., Boomsma, J. J., Ratnieks, F. L. W. Quantification of honey bee mating range and isolation in semi-isolated valleys in the Peak District National Park, England, as revealed by paternity analysis using DNA microsatellites. (submitted).

Medina, L. M., Martin, S. J., Ratnieks, F. L. W. Reproduction of Varroa destructor in drone brood cells of Africanized honey bees (Apis mellifera) in Yucatan, Mexico. (submitted)

Medina, L., Ratnieks, F. L. W. A scientific note on the occurence of hygienic behaviour in native British honey bees (Apis mellifera mellifera). (submitted)

Martin, S. J., Ratnieks, F. L. W., Jones, G. Dufour's gland and egg recognition in honey bees. (submitted)

Paxton, R. J., Ruhnke, H., Ratnieks, F. L. W., Quezada-Euan, J. J. G. Queen monoandry yet lack of worker maternity of males in the stingless bee Melipona beecheii (Apidae, Meliponini). (submitted)

Accepted for publication

Beekman, M., Seraphides N, Sumpter, D. J., Ratnieks, F. L. W. Patch use by honey bee colonies: does colony size matter? (accepted)

Chaline, N., Martin, S. J., Ratnieks, F. L. W. Absence of nepotism towards imprisoned young queens during swarming in the honey bee. Behavioral Ecology (accepted).

Ratnieks, F.L.W., Foster, K.R., Wenseleers, T. 2006. Conflict resolution in insect societies. Annual Review of Entomology, in press. (galleys, unrevised)

Wood, M. J., Ratnieks, F. L. W. Transfer of olfactory cues used in Vespula wasp recognition by honey bee guards. Apidologie (accepted).

Published

Jackson, D.E., Ratnieks, F.L.W. 2006. Primer: Communication in Ants. Current Biology, 16: R570-R574.

Tofilski, A., Ratnieks, F.L.W. 2006. Sand pile Formation in Dorymyrmex ants. Journal of Insect Behaviour, 18: 505-512.

117. Robinson EJH, Jackson DE, Holcombe M, Ratnieks FLW. 2005. 'No entry' signal in ant foraging. Nature, 438: 442. Supplementary Information

116. Ratnieks, F.L.W., Wenseleers, T. 2005. Policing Insect Societies. Science, 307: 54-56.

115. Wenseleers, T., Ratnieks, F.L.W., Ribeiro, M.F., Alves, D.A., Imperatriz-Fonseca, V.-L. Working-class royalty: bees beat the caste system. 2005. Biology Letters, 1: 125-128.

114. Wenseleers, T. Tofilski, A., Ratnieks, F.L.W. 2005. Queen and worker policing in the tree wasp Dolichovespula sylvestris. Behavioral Ecology & Sociobiology, 58: 80-86.

113. Wenseleers, T., Badcock, N.S., Erven, K., Tofilski, A., Nascimento, F.S., Hart, A.G., Burke, T.A., Archer, M.E., Ratnieks, F.L.W. A test of worker policing theory in an advanced eusocial wasp, Vespula rufa. Evolution, 59: 1306-1314.

112. Chaline, N., Martin, S. J., Ratnieks, F. L. W. 2004. Worker policing persists in a hopelessly queenless honey bee colony (Apis mellifera). Insectes Sociaux, 51: 113-116.

111. Chaline, N., Ratnieks, F. L. W., Raine, N. E., Badcock, N. S., Burke, T. 2004. Non-lethal sampling of honey bee, Apis mellifera, DNA using wing tips. Apidologie, 35: 311-318.

110. D'Ettorre, P., Heinze, J., Ratnieks, F. L. W. 2004. Worker policing by egg eating in the ponerine ant Pachycondyla inversa. Proceedings of the Royal Society of London Series B-Biological Sciences 271: 1427-1434.

109. Jackson, D. E., Holcombe, M., Ratnieks, F. L. W. 2004. Geometry gives polarity to ant pheromone trails. Nature 432: 907-909. Supplement1 Supplement2

108. Martin, S. J., Chaline, N., Oldroyd, B. P., Jones, G. R., Ratnieks, F. L. W. 2004. Egg marking pheromones of anarchistic worker honeybees (Apis mellifera). Behavioral Ecology 15: 839-844.

107. Martin, S.J., Jones, G. R., Chaline, N., Ratnieks F. L. W. 2004. Role of hydrocarbons in egg recognition in the honeybee. Physiological Entomology 29: 395-399.

106. Wenseleers T., Hart A. G., Ratnieks F.L.W. 2004. When resistance is useless: policing and the evolution of reproductive acquiescence in insect societies. American Naturalist 164: E154 -E167

105. Wenseleers, T., Hart, A. G., Ratnieks, F. L. W., Quezada-Euan, J. J. G. 2004. Queen execution and caste conflict in the stingless bee Melipona beecheii. Ethology, 110: 725-736.

104. Wenseleers, T., Helantera, H., Hart, A. G., Ratnieks, F. L. W. 2004. Worker reproduction and policing in insect societies: an ESS analysis. Journal of Evolutionary Biology 17: 1035-1047.

103. Wenseleers, T., Ratnieks, F. L. W. 2004. Tragedy of the commons in Melipona bees. Proceedings of the Royal Society of London Series B-Biological Sciences 271: S310-312. View talk on this (powerpoint)

102. Wood, M., Ratnieks, F.L.W. 2004. Olfactory cues and Vespula wasp recognition by honey bee guards. Apidologie 35: 461-468.

101. Beekman, M., Komdeur, J., Ratnieks, F. L. W. 2003. Reproductive conflicts in social animals - who has power. Trends in Ecology and Evolution, 18: 277-282.

100. Beekman, M., Ratnieks, F. L. W. 2003. Power over reproduction in social Hymenoptera. Philosophical Transactions of the Royal Society, 258: 1741-1753.

99. Chaline N, Arnold G, Papin C, et al. 2003. Patriline differences in emergency queen rearing in the honey bee, Apis mellifera. Insectes Sociaux 50: 234-236.

98. Gregson, A. M., Hart, A. G., Holcombe, M., Ratnieks, F. L. W. 2003. Partial nectar loads as a cause of multiple nectar transfer in the honey bee (Apis mellifera): a simulation model. Journal of Theoretical Biology 222: 1-8.

97. Hart, A. G., Bowtell, R. W., Kockenberger, W., Wenseleers, T., Ratnieks, F. L. W. 2003. Magnetic resonance imaging in entomology: a critical review. Journal of Insect Science, 3.5. 9 pages. Available online: insectscience.org/3.5

96. Jeanson, R., Ratnieks, F. L. W. & Deneubourg, J. L. 2003. Pheromone trail decay rates on different substrates in the Pharaoh's ant, Monomorium pharaonis. Physiological Entomology 28: 192-198.

95. Monnin T., Ratnieks F. L. W., Brandão C. R. F. 2003. Reproductive conflict in animal societies: hierarchy length increases with colony size in queenless ponerine ants. Behavioral Ecology & Sociobiology 54: 71-79.

94. Pirk, C. A. A., Neumann, P., Ratnieks, F. L. W. 2003. Cape honeybees police worker-laid eggs despite the absence of relatedness benefits. Behavioral Ecology, 14: 347-352.

93. Wenseleers, T., Ratnieks, F. L. W., Billen, J. 2003. Caste fate conflict in swarm-founding social Hymenoptera: an inclusive fitness analysis. Journal of Evolutionary Biology, 16: 647-658.

92. Beekman, M., Calis, J. N. M., Oldroyd, B. P., Ratnieks, F. L. W. 2002. When do honeybee guards reject their former nestmates after swarming? Insectes Sociaux 49: 56-61.

91. Beekman, M., Good, G., Allsopp, M. H., Radloff, S., Pirk, C. W. W., Ratnieks, F. L. W. 2002. A non-policing honey bee colony (Apis mellifera capensis). Naturwissenschaften 89: 479-482.

90. Beekman, M., Wossler, T. C., Martin, S. J., Ratnieks, F. L. W. 2002. Parasitic Cape honeybee workers (Apis mellifera capensis) are not given differential treatment by African guards (A. m. scuttelata). Insectes Sociaux 49: 216-220.

89. Chaline, N., Ratnieks, F. L. W., Burke, T. 2002. Anarchy in the UK: Detailed genetic analysis of worker reproduction in a naturally occurring British anarchistic honeybee, Apis mellifera, colony using DNA microsatellites. Molecular Ecology 11: 1795-1803.

88. Foster, K. R., Gulliver, J., Ratnieks, F. L. W. 2002. Worker policing in the European hornet Vespa crabro. Insectes Sociaux 49: 41-44.

87. Hart, A. G., Anderson, C., Ratnieks, F. L. W. 2002. Task partitioning in leafcutting ants. Acta Ethologica 5:1-11.

86. Hart, A. G. and Ratnieks, F. L. W. 2002. Task partitioned nectar transfer in stingless bees (Meliponini): work organisation in a phylogenetic context. Ecological Entomology 27: 163-168.

85. Hart, A. G. and Ratnieks, F. L. W. 2002. Waste management in the leafcutting ant Atta colombica. Behavioral Ecology 13: 224-231.

84. Martin, S. J., Beekman, M., Wossler, T. C., Ratnieks, F. L. W. 2002. Parasitic Cape honeybee workers, Apis mellifera capensis, evade policing. Nature 415: 163-165.

83. Martin, S. J., Jones, G. R., Châline, N., Middleton, H & Ratnieks, F. L. W. 2002. Reassessing the role of the honeybee (Apis mellifera) Dufour's gland in egg marking. Naturwissenschaften 89: 528-532.

82. Medina, L. M., Martin, S. J., Espinoza-Montaño, L., Ratnieks, F. L. W. 2002. Reproduction of Varroa destructor in worker brood of Africanized honey bees (Apis mellifera). Experimental and Applied Acarology 27: 79-88.

81. Monnin, T., Ratnieks, F. L. W., Jones, G. R., Beard, R. 2002. Pretender punishment induced by chemical signalling in a queenless ant. Nature 419: 61-65.

80. Oldroyd, B. P., Ratnieks, F. L. W., Wossler, T. C. 2002. Egg marking pheromones in honey-bees Apis mellifera. Behavioural Ecology and Sociobiology 51: 590-591.

79. Wattanachaiyingcharoen, W., Oldroyd, B. P., Good, G., Halling, L. Ratnieks, F. L. W., Wongsiri, S. 2002. Lack of worker reproduction in the giant honey bee Apis dorsata Fabricius. Insectes Sociaux 49: 80-85.

78. Barron, A. B., B. P. Oldroyd, and F. L. W. Ratnieks. 2001. Worker reproduction in honey-bees (Apis) and the anarchic syndrome: a review. Behavioral Ecology and Sociobiology 50:199-208.

77. Beekman, M., D. J. T. Sumpter, and F. L. W. Ratnieks. 2001. Phase transition between disordered and ordered foraging in Pharaoh's ants. Proceedings of the National Academy of Sciences of the United States of America 98:9703-9706.

76. Bourke, A. F. G., and F. L. W. Ratnieks. 2001. Kin-selected conflict in the bumble bee Bombus terrestris (Hymenoptera : Apidae). Proceedings of the Royal Society of London Series B-Biological Sciences 268:347-355.

75. Downs, S. G., F. L. W. Ratnieks, N. S. Badcock, and A. Mynott. 2001. Honeybee guards do not use food-derived odors to recognize non- nest mates: a test of the Odor Convergence hypothesis. Behavioral Ecology 12:47-50. 74. Foster, K. R., and F. L. W. Ratnieks 2001. The effect of sex allocation biasing on the evolution of worker policing. American Naturalist 158:615-623.

73. Foster, K. R., and F. L. W. Ratnieks. 2001. Convergent evolution of worker policing by egg eating in the honeybee and common wasp. Proceedings of the Royal Society of London Series B-Biological Sciences 268:169-174.

72. Foster, K. R., and F. L. W. Ratnieks. 2001. Paternity, reproduction and conflict in vespine wasps: a model system for testing kin selection predictions. Behavioral Ecology and Sociobiology 50:1-8.

71. Foster, K. R., F. L. W. Ratnieks, N. Gyllenstrand, and P. A. Thoren. 2001. Colony kin structure and male production in Dolichovespula wasps. Molecular Ecology 10:1003-1010.

70. Foster, K. R., T. Wenseleers, and F. L. W. Ratnieks. 2001. Spite: Hamilton's unproven theory. Annales Zoologici Fennici 38:229-238.

69. Hart, A. G., and F. L. W. Ratnieks. 2001. Leaf caching in the leafcutting ant Atta colombica: organizational shift, task partitioning and making the best of a bad job. Animal Behaviour 62:227-234.

68. Hart, A. G., and F. L. W. Ratnieks. 2001. Task partitioning, division of labour and nest compartmentalisation collectively isolate hazardous waste in the leafcutting ant Atta cephalotes. Behavioral Ecology and Sociobiology 49:387-392.

67. Hart, A. G., and F. L. W. Ratnieks. 2001. Why do honey-bee (Apis mellifera) foragers transfer nectar to several receivers? Information improvement through multiple sampling in a biological system. Behavioral Ecology and Sociobiology 49:244-250.

66. Miller, D. G., and F. L. W. Ratnieks. 2001. The timing of worker reproduction and breakdown of policing behaviour in queenless honey bee (Apis mellifera L.) societies. Insectes Sociaux 48:178-184.

65. Monnin, T., and F. L. W. Ratnieks. 2001. Policing in queenless ponerine ants. Behavioral Ecology and Sociobiology 50:97-108.

64. Neumann, P., C. W. W. Pirk, H. R. Hepburn, A. J. Solbrig, F. L. W. Ratnieks, P. J. Elzen, and J. R. Baxter. 2001. Social encapsulation of beetle parasites by Cape honeybee colonies (Apis mellifera capensis Esch.). Naturwissenschaften 88:214-216.

63. Oldroyd, B. P., L. A. Halling, G. Good, W. Wattanachaiyingcharoen, A. B. Barron, P. Nanork, S. Wongsiri, and F. L. W. Ratnieks. 2001. Worker policing and worker reproduction in Apis cerana. Behavioral Ecology and Sociobiology 50:371-377.

62. Oldroyd, B. P., T. C. Wossler, and F. L. W. Ratnieks. 2001. Regulation of ovary activation in worker honey-bees (Apis mellifera): larval signal production and adult response thresholds differ between anarchistic and wild-type bees. Behavioral Ecology and Sociobiology 50:366-370.

61. Ratnieks, F. L. W. 2001. Heirs and spares: caste conflict and excess queen production in Melipona bees. Behavioral Ecology and Sociobiology 5:467-473; DOI 10.1007/s002650100388. View talk on this (powerpoint)

60. Ratnieks, F. L. W., A. Lapish, and L. Tann. 2001. Honey bee guards, Apis mellifera, accept own subspecies non-nestmates more than other subspecies non-nestmates. Insectes Sociaux 48:287-288.

59. Ratnieks, F. L. W., T. Monnin, and K. R. Foster. 2001. Inclusive fitness theory: novel predictions and tests in eusocial Hymenoptera. Annales Zoologici Fennici 38:201-214.

58. Anderson, C., and F. L. W. Ratnieks. 2000. Task partitioning in insect societies: novel situations. Insectes Sociaux 47:198-199.

57. Beekman, M., and F. L. W. Ratnieks. 2000. Long-range foraging by the honey-bee, Apis mellifera L. Functional Ecology 14:490-496.

56. Bernasconi, G., F. L. W. Ratnieks, and E. Rand. 2000. Effect of "spraying" by fighting honey bee queens (Apis mellifera L.) on the temporal structure of fights. Insectes Sociaux 47:21-26.

55. Downs, S. G., and F. L. W. Ratnieks. 2000. Adaptive shifts in honey bee (Apis mellifera L.) guarding behavior support predictions of the acceptance threshold model. Behavioral Ecology 11:326-333.

54. Downs, S. G., F. L. W. Ratnieks, S. L. Jefferies, and H. E. Rigby. 2000. The role of floral oils in the nestmate recognition system of honey bees (Apis mellifera L.). Apidologie 31:357-365.

53. Foster, K. R., and F. L. W. Ratnieks. 2000. Facultative worker policing in a wasp. Nature 407:692-693.

52. Foster, K. R., F. L. W. Ratnieks, and A. F. Raybould. 2000. Do hornets have zombie workers? Molecular Ecology 9:735-742.

51. Foster, K. R., F. L. W. Ratnieks, and T. Wenseleers. 2000. Spite in social insects. Trends in Ecology and Evolution 15:469-470.

50. Hart, A. G., and F. L. W. Ratnieks. 2000. Leaf caching in Atta leafcutting ants: discrete cache formation through positive feedback. Animal Behaviour 59:587-591.

49. Oldroyd, B. P., and F. L. W. Ratnieks. 2000. Evolution of worker sterility in honey-bees (Apis mellifera): how anarchistic workers evade policing by laying eggs that have low removal rates. Behavioral Ecology and Sociobiology 47:268-273.

48. Ratnieks, F. L. W. 2000. Worker policing in the honey bee: basic facts and ideas. Insect Social Life 3:3-10.

47. Anderson, C., and F. L. W. Ratnieks. 1999. Task partitioning in insect societies. I. Effect of colony size on queueing delay and colony ergonomic efficiency. American Naturalist 154:521-535.

46. Anderson, C., and F. L. W. Ratnieks. 1999. Worker allocation in insect societies: coordination of nectar foragers and nectar receivers in the honey bee. Behavioral Ecology and Sociobiology 46:73-81.

45. Anderson, C., Ratnieks, F. L. W. 1999. Task partitioning in foraging: effect of colony size on queueing times and information reliability. Pages 31-50 in: Detrain C, Deneubourg J L, Pasteels J M (eds.). Information Processing in Social Insects. Birkhauser, Basel.

44. Bernasconi, G., Bigler, L., Hesse, M., Ratnieks, F. L. W. 1999. Characterization of queen specific components of the fluid released by fighting honey bee queens. Chemoecology 9: 161-167.

43. Bourke, A. F. G., and F. L. W. Ratnieks. 1999. Kin conflict over caste determination in social Hymenoptera. Behavioral Ecology and Sociobiology 46:287-297. View talk on this (powerpoint)

42. Downs, S. G., and F. L. W. Ratnieks. 1999. Recognition of conspecifics by honeybee guards uses nonheritable cues acquired in the adult stage. Animal Behaviour 58:643-648.

41. Foster, K. R., P. Seppa, F. L. W. Ratnieks, and P. A. Thorén. 1999. Low paternity in the hornet Vespa crabro indicates that multiple mating by queens is derived in vespine wasps. Behavioral Ecology and Sociobiology 46:252-257.

40. Monnin, T., and F. L. W. Ratnieks. 1999. Reproduction versus work in queenless ants: when to join a hierarchy of hopeful reproductives? Behavioral Ecology and Sociobiology 46:413-422.

39. Ratnieks, F. L. W., Anderson, C. 1999. Task partitioning in insect societies: natural history, ergonomics, and information. Actes Colloq. Insectes Sociaux. 12: 1-10.

38. Ratnieks, F. L. W., and C. Anderson. 1999. Task partitioning in insect societies. Insectes Sociaux 46:95-108.

37. Ratnieks, F. L. W., and C. Anderson. 1999. Task partitioning in insect societies. II. Use of queueing delay information in recruitment. American Naturalist 154:536-548.

36. Ratnieks, F. L. W., Keller, L. 1998. Queen control of egg fertilization in the honey bee. Behavioral Ecology and Sociobiology 44: 57-61.

35. Sundström, L., Ratnieks, F. L. W. 1998. Sex ratio conflicts, mating frequency and queen fitness in the ant Formica truncorum. Behavioral Ecology 9:116-121.

34. Ratnieks, F. L. W., Boomsma, J. J. 1997. On the robustness of split sex ratio predictions in monogynous social Hymenoptera. Journal of Theoretical Biology 185: 423-439.

33. Ratnieks, F. L. W. 1996. Evolution of unstable and stable biparental care. Behavioral Ecology : 490-493.

32. Ratnieks, F. L. W., Visscher, P. K. 1996. Living with the Africanized bee...Sinaloao beekeepers adapt pollination to Africanized bees. California Agriculture. 50 (4) 24-28.

31. Boomsma, J. J., Ratnieks, F. L. W. 1996. Paternity in eusocial hymenoptera. Philosophical Transactions of the Royal Society of London, B. 351: 947-975.

30. Ratnieks, F. L. W., Vetter, R., Visscher, P. K. 1996. A polygynous overwintered nest of Vespula pensylvanica from southern California. Insectes Sociaux 43: 401-410.

29. Ratnieks, F. L. W. 1995. Evidence for a queen-produced egg-marking pheromone and its use in worker policing in the honey bee. Journal of Apicultural Research 34: 31-37.

28. Ratnieks, F. L. W., Boomsma, J. J. 1995. Facultative sex allocation by workers and the evolution of polyandry by queens in social Hymenoptera. American Naturalist 145: 969-993.

27. Ratnieks, F. L. W., Visscher, P. K, Vetter, R. 1995. Treating American foulbrood using gamma radiation: a case history. American Bee Journal 135: 557-561.

26. Ratnieks, F. L. W., Miller, D. G. 1993. Natural history of polygynous Vespula vulgaris colonies in California. Psyche 100: 43-50.

25. Ratnieks, F. L. W., Miller, D. G. 1993. Division of honey bee drones during swarming. Animal Behaviour 46: 803-805.

24. Morse, R. A., Layne, J. N., Visscher, P. K., Ratnieks, F. 1993. Selection of nest cavity volume and entrance size by honey bees in Florida. Florida Scientist 56: 163-167.

23. Ratnieks, F. L. W. 1993. Egg-laying, egg-removal, and ovary development by workers in queenright honey bee colonies. Behavioral Ecology and Sociobiology 32: 191-198.

22. Reeve, H. K., Ratnieks, F. L. W. 1993. Queen conflict in polygynous societies: mutual tolerance and reproductive skew. In: Keller, L. (ed). Queen number and sociality in insects. Oxford University Press, Oxford.

21. Ratnieks, F. L. W., Reeve, H. K. 1992. The structure of conflict in hymenopteran societies: processes that reduce conflict in advanced eusocial species. Journal of Theoretical Biology 158: 33-65.

20. Getz, W. M., Kaitala, V., Ratnieks, F. L. W. 1992. Invasion of sibmating genes in diploid and haplodiploid populations. Evolutionary Ecology 6: 312-330.

19. Ratnieks, F. L. W. 1992. American foulbrood: A review of an important honey bee disease. Bee World 73: 177-191.

18. Ratnieks, F. L. W. 1991. Evolution of discriminatory aggression in marine invertebrates. Journal of Theoretical Biology 152: 557-565.

17. Emlen, S. T., Reeve, H. K., Sherman, P. W., Wrege, P. H., Ratnieks, F. L. W., Shellman-Reeve, J. 1991. Adaptive versus non-adaptive explanations of behavior: the case of alloparental helping. American Naturalist 138: 259-270.

16. Ratnieks, F. L. W., Piery, M., Cuadriello, I. 1991. The natural nest of the Africanized honey bee (Hymenoptera, Apidae) near Tapachula, Chiapas, Mexico. Canadian Entomologist 123: 353-359.

15. Ratnieks, F. L. W. 1991. Africanized bees: Natural selection for colonizing ability. Pages 119-135. In: Spivak, M., Breed, M. D., Fletcher, D. J. C. (eds) The Africanized honey bee. Westview Press, Boulder, CO.

14. Ratnieks, F. L. W. 1991. Facultative sex allocation biasing by workers in social Hymenoptera. Evolution 45: 281-292.

13. Ratnieks, F. L. W. 1991. The evolution of genetic cue diversity in social Hymenoptera. American Naturalist 137: 202-226.

12. Ratnieks, F. L. W., Reeve, H. K. 1991. The evolution of queen-rearing nepotism in social Hymenoptera: effects of discrimination costs in swarming species. Journal of Evolutionary Biology 4: 93-115.

11. Ratnieks, F. L. W. 1990. The evolution of polyandry by highly eusocial bee queens: The importance of progressive versus mass provisioning and the timing of removal of diploid males. Behavioral Ecology & Sociobiology 26: 342-348.

10. Ratnieks, F. L. W. 1990. Assessment of queen mating frequency by workers in social Hymenoptera. Journal of Theoretical Biology 142: 87-94.

9. Morse, R. A., Camazine, S., Ferracane, M., Minacci, P., Ratnieks, F. L. W., Spielholz, J., Underwood, B.A. 1990. The population density of feral honey bee colonies (Hymenoptera, Apidae) in the city of Oswego, New York. Journal of Economic Entomology 83: 81-83.

8. Ratnieks, F. L. W., Visscher, P. K. 1989. Worker policing in the honey bee. Nature 342: 796-797.

7. Ratnieks, F. L. W., Nowakowski, J. 1989. Honey bee swarms accept bait hives contaminated with American foulbrood disease. Ecological Entomology 14: 475-478.

6. Mumme, R. L., Koenig, W. G., Ratnieks, F. L. W. 1989. Helping behaviour, reproductive value, and the future component of indirect fitness. Animal Behaviour 38: 331-343.

5. Ratnieks, F. L. W. 1989. Sterilization of American foulbrood contaminated hive equipment using gamma irradiation. American Bee Journal 129: 191-193.

4. Kukuk, P. F., Eickwort, G. C., Raveret-Richter, M., Alexander, B., Gibson, R., Morse, R. A., Ratnieks, F. L. W. 1989. The importance of the sting in the evolution of sociality in the Hymenoptera. Annals of the Entomological Society of America 82: 1-5.

3. Ratnieks, F. L. W. 1988. Reproductive harmony via mutual policing by workers in eusocial Hymenoptera. American Naturalist 132: 217-236.

2. Ratnieks, F. L. W. 1988. Improved bait hives. American Bee Journal 128: 125-7.

1. Robinson, G. E., Ratnieks, F. L. W. 1987. Induction of premature foraging by orally and topically applied juvenile hormone analogs in the honey bee (Apis mellifera L.). Journal of Economic Entomology 80: 784-787.

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articles in conference proceedings

21. Ratnieks, F. L. W. 2002. Parasitism of African honey bee colonies by cape honey bee workers. Pages 81-83 in Garófalo, C. (ed.) V Encontro sobre abelhas. Universidade de São Paulo, Ribeirão Preto.

20. Ratnieks, F. L. W. 2002. Cooperation through coercion: policing of male production and female caste fate in honey bees and stingless bee. Pages 10-14 in Garófalo, C. (ed.) V Encontro sobre abelhas. Universidade de São Paulo, Ribeirão Preto.

19. Ratnieks, F. L. W., Monnin, T. 2002. Dominance hierarchies and conflict: insights from inclusive fitness theory. Page 74 in: Proceedings XIV International Congress of IUSSI, Sapporo, Japan.

18. Wossler, T. C., Oldroyd, B. P., Ratnieks, F. L. W. 2002. Reproductive anarchy in the beehive: the role of pheromones. Page 213 in: Proceedings XIV International Congress of IUSSI, Sapporo, Japan.

17. Martin, S. J., Ratnieks, F. L. W., Jones, G. R.. 2002. How do honeybee workers discriminate between worker-laid and queen-laid eggs? Page 124 in: Proceedings XIV International Congress of IUSSI, Sapporo, Japan.

16. D'Ettorre, P., Heinze, J., Ratnieks, F. L. W. 2002. Policing by egg eating in the ponerine ant Pachycondyla inversa. Page 77 in: Proceedings XIV International Congress of IUSSI, Sapporo, Japan.

15. Monnin, T., Ratnieks, F. L. W., Jones, G. R., Beard, R. 2002. Page 76 in: Proceedings XIV International Congress of IUSSI, Sapporo, Japan.

14. Beekman, M, Ratnieks, F. L. W. 2002. Power over reproduction in eusocial Hymenoptera. Page 74 in: Proceedings XIV International Congress of IUSSI, Sapporo, Japan.

13. The Mathematical Modelling of Social Behaviour. EU 'INSECTS' TMR meeting, Great Hucklow, UK, 14-18 Sept. 2001, 113 pages.

12. Ratnieks, F. L. W. 2001. Por que las abejas Melipona crian reinas en exceso? El conflicto de castas y el poder de las larvas sobre su propio destino (Why do Melipona bees rear excess queens? Caste fate conflict and the power of larvae over their fate). Pages 102-103 in: Quezada-Euán, J., May-Itzá, W., Moo-Valle, H., Medina, J. (eds) II Seminario Mexicano sobre abejas sin aguijón.

11. Paxton, R., Ruhnke, H., Shah, M., Bego, L. R., Quezada-Euán, J. J. G, Ratnieks, F. L. W.2001. Social evolution in stingless bees: are the workers or is the queen in control of male parentage? Pages 108-107 in: Quezada-Euán, J., May-Itzá, W., Moo-Valle, H., Medina, J. (eds) II Seminario Mexicano sobre abejas sin aguijón.

10. Ratnieks, F. L. W. 2001. Our best theory: Novel predictions and tests on socila insect biology arising from W. D. Hamilton;s theory of inclusive fitness 35 years on. Page 16 in: Menzel, R., Rademacher, E. Proceedings of the 2001 Berlin meeting of the European sections of IUSSI (International Union for the Study of Social Insects).

9. Ratnieks, F. L. W. 2001. Conflict and caste determination. Page 100a in: Menzel, R., Rademacher, E. Proceedings of the 2001 Berlin meeting of the European sections of IUSSI (International Union for the Study of Social Insects).

8. Ruhnke, H., Shah, M., Mecke, R., Bego, L. R., Quezada-Euán, J., Ratnieks, F. L. W., Paxton, R. J. 2001. Kin conflict and the parentage of males in stingless bees. Page 58 in: Menzel, R., Rademacher, E. Proceedings of the 2001 Berlin meeting of the European sections of IUSSI (International Union for the Study of Social Insects).

7. Ratnieks, F. L. W. 1998. Information sources in reproductive manipulation. Page 387 in: Schwarz, M. P., Hogendoorn, K. (eds) Proceedings of the 13th International Congress of IUSSI. IUSSI.

6. Monnin, T., Peeters, C., Ratnieks, F. L. W. 1998. "Queen" policing and worker policing in the queenless ant Dinoponera quadriceps. Page 323 in: Schwarz, M. P., Hogendoorn, K. (eds) Proceedings of the 13th International Congress of IUSSI. IUSSI.

5. Foster, K. R., Seppä, P., Ratnieks, F. L. W., Thorén, P. 1998. Low paternity and queen-worker conflict in the hornet. Page 168 in: Schwarz, M. P., Hogendoorn, K. (eds) Proceedings of the 13th International Congress of IUSSI. IUSSI.

4. Ratnieks, F. L. W. 1996. Worker reproduction and worker policing: how the balance is set. p. 124, Abstracts of: Fifth International Conference of Systematic and Evolutionary Biology.

3. Ratnieks, F. L. W. 1994. Worker policing in honey bees. page 582 in: Lenoir, A., Arnold, G., Lepage, M. (eds) Les Insectes Sociaux. (Proceedings of the 12th International Congress of IUSSI). Université Paris Nord, Paris.

2. Ratnieks, F. L. W. 1992. Evidence for an egg-marking pheromone in the honey bee. American Bee Journal 132: 813.

1. Ratnieks, F. L. W. 1987. The adaptive significance of nest site selection by honey bee swarms: calculation of reproductive value of swarms using colony life table data. Page 617 in: J. Eder and H. Rembold (eds) Chemistry and biology of social insects. Verlag J. Peperny, Munich.

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talks, courses and lecture notes

For dedicated web pages for the courses APS209, 301 and 323 see

butt3.gif (904 bytes) APS 209 Animal Behaviour butt3.gif (904 bytes) APS 301 Conflict & Cooperation butt3.gif (904 bytes) APS 323 Social Insects

Ratnieks, F.L.W. 2004. Conflict resolution by policing in insect societies. Talk at the Wissenschaftkolleg zu Berlin (Institute for Advanced Study), 21 Dec. 2004. View lecture (powerpoint)

Wenseleers, T. 2002. Wolbachia : microbial manipulator of insect reproduction. Guest lecture in the Selfishness & Altruism course. View lecture (powerpoint)

The Mathematical Modelling of Social Behaviour. EU 'INSECTS' TMR meeting, Great Hucklow, UK, 14-18 Sept. 2001, 113 pages.

Wenseleers, T. 2001. An introduction to the evolutionary modelling of conflict. In: "The Mathematical Modelling of Social Behaviour" (T. Wenseleers & F.L.W. Ratnieks, eds.), p. 21-53, EU INSECTS training workshop, Great Hucklow, UK, 14-18 Sept. 2001, 113 pages. Overheads Mathematica notebooks

Wenseleers, T. and Ratnieks, F. L. W. 2002. Actual and potential tragedies: conflict over caste fate in social insects. Seminar, University of Sheffield. View talk (powerpoint)

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extension and popular articles on beekeeping and social insects

52. Robinson, E.J.H., Tofilski, A., Ratnieks, F.L.W. 2004 Upper Dearne Woodlands Wood Ant Survey.

51. Ratnieks, F. L. W., Jensen, A. 2003. Evaluation of queen mating isolation in the Peak district, Sheffield, by means of DNA microsatellites. Bee Improvement, 15: 10-12.

50. Big Business: Bijenkolonies en oliemaatschappijen. Maandblad van de Vlaamse Imkerbond. December 2003, p. 304-306. Translation and Adaptation in Dutch of F. Ratnieks (2002) "Big Business - Honeybee colonies and oil companies" by A. Schotanus and T. Wenseleers.

49. Conflict in het bijenvolk. Maandblad van de Vlaamse Imkerbond. Juli-Augustus 2003, p. 186-188. Translation and Adaptation in Dutch of "F. Ratnieks (2002) Conflict in the Bee Hive: Worker Reproduction and Worker Policing" by A. Schotanus and T. Wenseleers.

48. Ratnieks, F. L. W. 2003. Punishment, policing and conflict in Dinosaur ants (pamphlet)

47. Ratnieks, F. L. W. 2003. A farming bee. The Beekeepers Quarterly (in press).

46. Ratnieks, F. L. W. 2003. The BABE Project: Beekeeping and Apis Biodiversity in Europe. BBKA News (in press)

45. Ratnieks, F. L. W. 2002. Laboratory of Apiculture and Social Insects: The story so far. The Beekeepers Quarterly 71: 37-39.

44. Ratnieks, F. L. W., Palmer, K. 2002. BABE research in Hope Valley and Edale, Derbyshire, Summer 2002. Bee Improvement 12: 22-23.

43. Ratnieks, F. L. W. 2002. Comb honey. Pamphlet.

42. Ratnieks, F. L. W. 2002. Conflict in the bee hive: worker reproduction and worker policing. The Beekeepers Quarterly 70: 16-17.

41. Ratnieks, F. L. W., Hart, A. G. 2002. Garbage dispoal in leafcutter ants, Atta. Pamphlet.

40. Ratnieks, F. L. W. 2002. The honey bee, Apis mellifera. Pamphlet.

39. Ratnieks, F. L. W. 2002. The black bee, Apis mellifera mellifera. Pamphlet.

38. Ratnieks, F. L. W. 2002. Beekeeping. Pamphlet.

37. Ratnieks, F. L. W. 2002. How bees make honey. Pamphlet.

36. Ratnieks, F. L. W. 2002. Big business: honey bee colonies and oil companies. The Beekeepers Quarterly 69: 22-24.

35. Ratnieks, F. L. W. 2001. Are you being served? Supermarkets and bee hives. The Beekeepers Quarterly 67: 26-27.

34. Ratnieks, F. L. W. 2001. The last word: Bee lines 1. New Scientist 2301 (28 July 2001): 137.

33. Ratnieks, F. L. W. 2001. The last word: Bee lines 2. New Scientist 2303 (11 August 2001): 93.

32. Ratnieks, F. L. W. 2001. Prisons in the bee hive. The Beekeepers Quarterly 66: 15-16.

31. Ratnieks, F. L. W. 2001. Laboratory of Apiculture and Social Insects. The Beekeepers Quarterly 65: 9-10.

30. Ratnieks, F. L. W., Hart, A. G. 2001. Melipona beecheii beekeeping in the Yucatan. Bee Improvement

29. Ratnieks, F. L. W., Hart, A. G. 2001. Meliponinae bees: the other highly social bees. Bee Improvement

28. Ratnieks, F. L. W. 2001. Sociality and eusociality in insects. Bee Improvement 8: 20-21

27. Ratnieks, F. L. W. 2000. BABE: Beekeeping and Apis biodiversity in Europe. Bee Improvement 7: 13

26. Ratnieks, F. L. W. 2000. Why do researchers study the honey bee? Bee Improvement 7: 4-5

25. Ratnieks, F. L. W. 2000. How far do bees forage. Bee Improvement 6: 10-11.

24. Ratnieks, F. L. W., Pedersen, B. V. 1999. Phylogeny of British black bees. Bee Improvement 5: 6-7

23. Ratnieks, F. L. W. 1999. The honey bee and beekeeping. pamphlet

22. Ratnieks, F. L. W. 1999. Honey bee (Apis mellifera). pamphlet

21. Ratnieks, F. L. W. 1999. Dinosaur ants (Dinoponera). pamphlet

20. Ratnieks, F. L. W. 1999. Leafcutter ants (Atta). pamphlet

19. Ratnieks, F. L. W. 1999. Naked mole rat (Heterocephalus glaber). pamphlet

18. Ratnieks, F. L. W. 1999. Leaf-cutter and dinosaur ants. Pamphlet produced as part of Royal Society & British Association Millenium Award "Bringing bee biology and behaviour to beekeepers and the public".

17. Ratnieks, F. L. W. 1999. Tasmanian black bees. Bee Improvement 4: 5-6.

16. Ratnieks, F. L. W. 1998. Why breeding bees is different. Bee Improvement 2: 6-7.

15. Ratnieks, F. L. W. 1996. Varroa: What to do. Beekeepers News

14. Ratnieks, F. L. W. 1996. Langstroth, WBC, or National hive—the bees don't mind but you might! Beekeepers News

13. Ratnieks, F. L. W. 1996. Do you have nasty (defensive) bees? Then do something. Beekeepers News.

12. Ratnieks, F. L. W. 1996. Beekeeping is not cruel to bees. Beekeepers News

11. Ratnieks, F. L. W. 1994. Cox Honeyland: an innovative honey business from Utah. Gleanings in Bee Culture.

10. Ratnieks, F. L. W. 1992. Conflict and cooperation in the bee hive. Proceedings of the Western Apicultural Society 25-29.

9. Ferracane, M., Ratnieks, F. L. W. 1989. Africanized bees and good neighbors. Keeping bees in urban areas. Am. Bee J. 129: 117-121.

8. Ratnieks, F. L. W. 1989. Keep it simple. A simplified Langstroth hive.Gleanings in Bee Culture 117: 41-43.

7. Ratnieks, F. L. W., Nowogrodzki, R. 1988. Small Scale Queen Rearing. Cornell University Cooperative Extension. Information Bulletin 209.

6. Ratnieks, F. L. W. 1988. Disease update: The spread of American foulbrood within a honey bee colony. Gleanings in Bee Culture 116:

5. Ratnieks, F. L. W., Nolan, T.A. 1988. New York Beekeepers and their knowledge and need for information on brood diseases. Am. Bee J. 128: 761-764.

4. Ratnieks, F. L. W.1987. Bee space. Gleanings in Bee Culture 115: 573-5, 7.

3. Ratnieks, F. L. W.1987. Fall feeding. Gleanings in Bee Culture 115: 588-590.

2. Ratnieks, F. L. W., Nowogrodzki, R. 1987. The hobby beekeeper and American foulbrood: How to wisely use antibiotics as a preventive. Information bulletin of the New York State Apiary Inspection Program.

1. Ratnieks, F. L. W. 1985. What is a mite? Gleanings in Bee Culture 113: 483-484.

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book reviews

4. Ratnieks, F. L. W. 2003. review of Weiss, M. 2002. The little book of bees. Copernicus, New York. Antenna (Bulletin of the Royal Entomological Society of London)

3. Ratnieks, F. L. W. 1999. Review of: Archer, Michael E. 1998. Threatened wasps, ants and bees (Hymenoptera: Aculeata) in Watsonian Yorkshire. Page 43. Yorkshire Philosophical Society, Annual Report 1998.

2. Ratnieks, F. L. W. 1996. Review of "Natural history and evolution of paper wasps" edited by S. Turrilazzi & M. J. West-Eberhard. Oxford University Press. (reviewed for Animal Behaviour)

1. Ratnieks, F. L. W. Review of "The wisdom of the hive" by T. D. Seeley. Harvard University Press (reviewed for BeeBiz)

butt3.gif (904 bytes) media

Ratnieks, F. L. W. 1998. Dancing for their supper: the dance language and foraging ecology of the honey bee. Educational Video, Sheffield University Television.

butt3.gif (904 bytes) articles featuring our research

Whitfield, J. 2002. The police state. Nature 416: 782-784. Nature magazine news article featuring our research on bee policing

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